Elijah, Judah, Micah, and Josiah have been hired by the Lethal Company to extract scrap, in hopes of hitting quota, whilst avoiding deadly monsters. Meanwhile, a hooded figure (revealed to be Shiloh) is caught in the relics room.
The group travel to a mysterious realm and suffer a strange case on amnesia. They must recollect their memory and complete their original mission, whilst narrowly avoiding monsters. Meanwhile, Elijah realizes what Shiloh has been up to.
After Dry Bones Bowser challenges Princess Peach to a grand prix race, it's up to Shiloh and Bros to win the race and save the Mushroom Kingdom. But who summoned Dry Bone Bowser in the first place?