- A chainsaw-wielding George Washington teams with beer-loving bro Sam Adams to take down the Brits in a tongue-in-cheek riff on the American Revolution.
- In January 1776, just after signing the Declaration of Independence, its signatories are murdered by Benedict Arnold (Andy Samberg), who betrayed them and turned them over to the British side. He then goes to the Ford's Theater where George Washington (Channing Tatum) and his childhood best friend, Abraham "Abe" Lincoln (Will Forte), watch a play (Abe argues over the high prices of merchandise at the theatre ($5 for the item, and $30 in taxes)). Arnold turns into a werewolf (& tears up the signed declaration of independence) and kills Lincoln, who, before his death, asks Washington to bring the American Revolutionary War to an end and to name the recovered country "America."
Arnold leaves Washington alive as he is considered to be a party boy and not a serious leader like Lincoln. But Washington fights back with a pair of chainsaws and Arnold only escapes with the help of the transporter (Zebbie Gillese).
At Lincoln's funeral, Washington (who is the inventor of peanut butter) meets Martha Dandridge (Judy Greer), who urges him to continue the revolution after the two have sexual intercourse. In order to build a strong team, Washington teams up with party animal Samuel "Uncle Sam" Adams (Jason Mantzoukas) (who plays in Martha's adult kickball league. He runs a frat house and is known as a trouble maker. Sam hates the Brits as they can't handle the beer of the colonists, which is made from rotten grass), female Chinese scientist Thomas Edison (Olivia Munn) (who was being burnt at the stake for creating and testing hypothesis. She uses her science gadgets to kill the British soldiers and escape. Sam is outraged when Thomas turns out to be an immigrant chick. She also says she is not a sorcerer and that she is a scientist. She is willing to help Washington defeat the Brits as the Brits are trying to kill her), Native American hunter Geronimo (Raoul Max Trujillo) (he is the best tracker in the world. He is suspicious of Washington but agrees to help topple the Brits when he promises to return the land to the native Indians when they drive King James out), and equestrian champion Paul Revere (Bobby Moynihan) (Sam introduces Washington to Paul at a quarter mile horse-racing track. The illegal race is stopped by the "fun police", a moniker for the Brits).
They manage to track down Arnold at the Vietnam bar. Although the enemy has fled, they conclude that the events in Vietnam were not a failure (Arnold bites Geronimo and he has to chop off his arm to stop turning into a werewolf himself) (They give chase to Arnold on Clyde (Revere's horse), but Arnold manages to escape on a Jet Ski in the Boston River). Washington is crestfallen and wants to quit, but Martha tells him to quit only when the goals are realized. They ask for help from master blacksmith John Henry (Killer Mike), who is to make a silver bullet for them to kill Arnold. He asks Washington to bring him 1 metric ton of silver to cast 1 silver bullet.
To get the silver, they go to Boston to steal the supply of silver spoons from the British ship Titanic. Onboard the ship, they learn that the British are bathing people in tea to come over to their side. They also discover plans of a secret British super-weapon but fail to retrieve them because they sink the ship prematurely. Sam then proposes to the others they will say the Titanic had hit an iceberg and that they were never there.
Arnold kidnaps Martha, who is to become the future wife of King James (Simon Pegg). King James is working with Arnold to create a diabolical war machine and he intends to lure Washington into war and commit all his forces, so he can defeat the rebellion for all time. Washington manages to find out about a Gettysburg address (1 Merica St, which he guesses when he equals A to 1 in the name that Abe gave him) where Arnold meets the king at a secret meeting. The place turns out to be an ambush in which Clyde (Carlos Alazraqui), Revere's horse, dies, sacrificing himself by running away with the bomb that Arnold planted for the rebel group. King James appears at the Gettysburg address in the form of a hologram and offers Washington a battle the next morning at Attrition field, to which Washington agrees (He has no choice as King James threatens to sexually assault Martha).
Soon after, Arnold kills the king to take over the world himself. The next day, the British army stands up against the American army hastily assembled by Washington. On the side of the British, there are, among others, Elizabeth Tower in the form of a Transformer, London buses in the form of AT-ATs and Arnold flying in a crown-shaped ship. The side of the Americans consists of Paul Bunyan and Babe the Blue Ox, as well as Indians, Mexicans, African Americans, Asians, and Arabs, and Revere who had fused himself with what was left of Clyde with Nano-cybernetic technology by Clara Barton (Megan Leahy).
During the war, the British were trying to turn all revolutionaries British by tainting the clouds with tea and causing a tea rain. Edison, thanks to the power of science, manages to use the British weapons against them, turning tea into beer. When the beer rain falls, the British turn into Americans, and it also cures Martha from being one of the British. A final confrontation occurs against Arnold, who becomes a Mega Wolf until he is eventually defeated with a silver bullet from Henry, who kills him with America's favorite pastime.
Six months later, on July 4, 1776, Washington inaugurates the grand opening of America at the monument of his name. As George plans I.M.Pei (Dave Callaham) to build another memorial for his best friend to stare at his monument, the spirit of Abe comes back. It gives him the Declaration, repaired with scotch tape, while Samuel becomes George's uncle for his newborn son Denzel. The inauguration is disrupted as the crowd engages in a fight with each other, arguing over racism, abolitionism, women's rights, giving back the land to the natives, right to keep and bear arms, same-sex marriages, religious beliefs, free health care, and fair trials, making Washington nervous about the future of his country.
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By what name was America: The Motion Picture (2021) officially released in Japan in Japanese?
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