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- An invisible and mysterious force field descends upon the small town of Chester's Mill, Maine, USA, trapping residents inside, cut off from the rest of civilization. The trapped townspeople must discover the secrets and purpose of the "dome" or "sphere" and its origins, while coming to learn more than they ever knew about each other and animals too.
- Centers on an ultra-competitive surgeon whose life is changed forever when his ex-wife dies and begins teaching him what life is all about from the here-after.
- "THE LIGHT," IS A SPECIAL GLOBAL VIDEO TRIBUTE TO MEMORIALIZE THE ORLANDO PULSE NIGHTCLUB VICTIMS Created by Shawn Balentine, Hollywood's Openly Gay, Out Stuntman/Stunt Coordinator.
- After the ghost of Michael's ex-wife visits him from the afterlife and asks him to help continue running her low income clinic, he finds himself rushing back and forth between taking care of his own patients at Holt Neuro and her less fortunate patients at the clinic.
- While interviewing doctors to run the clinic, Michael helps a man who passed out, but refuses care due to lack of insurance. Meanwhile, as Michael evaluates a woman, he notices lesions on her brain that require immediate surgery, even as he doesn't know the exact diagnosis.
- Michael and Kate are forced to work together to try and save a patient with a deadly virus. Meanwhile, at Holt Neuro, as Michael treats a nervous bride-to-be who is experiencing headaches, he notices her mother is exhibiting symptoms of a serious condition.
- While at the clinic, Michael runs into a former football star that he performed successful surgery on years ago, and is now a shell of his former self, living on the streets. At Holt Neuro, Michael examines a patient who suffers from post-traumatic stress disorder.
- Michael works double duty trying to save a pregnant personal friend he recently diagnosed with a brain tumor and a little boy with sickle cell disease.
- Michael must decide if it is worth risking his practice to save a patient's life. Meanwhile, he courts Evan "E-Mo" Morris, a neuropsychiatrist to join his practice after E-Mo determines a patient's diagnosis by reading her body language.
- When Michael treats a patient who is hearing voices, he turns to Anton for help. Meanwhile, Kate's first day at the clinic doesn't go well.
- When Michael finds out a patient with a malignant brain tumor committed a horrible crime, he struggles with whether to save the man's life. Meanwhile, a female firefighter who appears to be disoriented is brought to Holt Neuro for evaluation.
- While tending to patients at the clinic during a blizzard, Michael shows signs of a severe condition that requires immediate surgery.
- Dr. Barnes and Dr. Sykora save a man on a bus who turns out to have a DNR order. Meanwhile, Rita's son has a motorcycle accident that reveals an existing medical condition.
- Michael and E-Mo disagree about how to treat a patient with a brain tumor. Meanwhile, a teenage girl gives birth at the clinic and immediately runs out, leaving the baby behind.
- Michael saves a man from drowning and learns his identity is a mystery.
- When a spoiled pop star comes to Holt Neuro experiencing unusual symptoms, and a young girl at the Clinica exhibits similar symptoms on the same day, it forces both teams of doctors to try and piece together their common link.
- When Michael's high school sweetheart comes to Holt Neuro with an irreversible condition, he will stop at nothing to try and save her, until Anna intervenes. Meanwhile, Zeke deals with the news that his best friend's cancer has returned.
- A distraught woman takes the Clinica staff hostage after Michael testifies for a former patient who he believed suffered uncontrollable urges due to a brain tumor.
- Michael learns that a patient at the clinic has a special connection to Anna. (Frances Fisher guest stars)
- In Chester's Mill, Maine, a strange occurrence locks the town in under an invisible dome-like surface.
- Following Duke's death, his house catches fire, forcing the town to rally together to form a bucket brigade. Elsewhere, Junior ambushes Barbie, while Dodee and Phil pick up a military frequency.
- After a Deputy is killed, Big Jim organizes a search party to hunt down the culprit. Meanwhile, Joe and Norrie continue to experience strange seizures.
- The town suffers from an outbreak of meningitis. Julia learns that Phil and Barbie are connected to her missing husband. Angie struggles to escape the bomb shelter.
- When friends and family from outside the dome come to visit, Barbie learns the military plan to destroy the dome with a missile.
- After Julia and Dodee speculate that the source of the radio signal jam could be the answer to the dome's origin, they are led to Joe and Norrie. Meanwhile, Angie seeks help after escaping from Junior.
- A friend of Julia's goes into labour after touching the dome. Big Jim kicks out Junior. Linda seeks out the Dundee brothers after Rose's murder. Joe and Norrie find a second mini-dome that holds a mysterious egg inside it.
- Norrie grieves over her mom's death. Joe shows Julia the mini-dome. Big Jim makes a plan of action when Ollie takes control of food and water.
- Julia takes Barbie to see the mini-dome and the egg, but they have both disappeared. A former acquaintance of Big Jim and Barbie makes a re-appearance. Angie starts to experience the same seizures as Joe and Norrie.
- Maxine continues to manipulate Barbie and Big Jim. Linda and Julia investigate Duke's involvement with previous criminal activity in Chester's Mill. Meanwhile, Angie realizes Junior is the "fourth hand".
- After Junior abandons Angie, Joe and Norrie, the weather turns nasty. Julia is shot by Maxine. Big Jim convinces Linda his stash of propane is what's saving the town and the biggest threat they face is from Barbie.
- Big Jim rallies the town against Barbie after framing him for the murders and the shooting of Julia. Joe and Norrie move the mini-dome to Ben's house but it soon begins to act strangely.
- As Barbie's fate is sealed by Big Jim, the dome blackens and the identity of the Monarch is revealed.
- As Barbie's fate lies in Big Jim's hands, the dome presents a new threat when it becomes magnetized. Meanwhile, Julia seeks out the help of a stranger to save the life of a mysterious girl.
- As the news of Angie's murder spreads, Rebecca warns Big Jim of a caterpillar infestation that could infect the town's remaining crops.
- While Julia and Sam attempt to stop Big Jim and Rebecca's plans for population control, Barbie, Joe and Norrie try to jog Melanie's memory, and Junior turns to Lyle for answers.
- Julia takes over as leader of Chester's Mill after the town becomes divided in the wake of Big Jim and Rebecca's plans for population control. Meanwhile, Joe and Norrie continue to help Melanie search for more clues about her identity.
- When Barbie and Sam investigate the mysterious tunnel beneath the high school, a cave-in severs their path back to Chester's Mill. Meanwhile, Big Jim deals with a dust storm.
- Barbie, Sam, Pauline, Lyle and Hunter try to evade Don and his forces and find a way back to Chester's Mill, while Junior and Melanie hide the egg from Julia and Big Jim.
- As resources dwindle, Big Jim holds a census in order to forecast how long Chester's Mill can continue to exist under the dome, while chaos ensues when a rainstorm turns into acid rain.
- After finding themselves outside the dome, Barbie turns to his estranged father for help, and Sam reunites with Pauline.
- Barbie realizes Don knows more about the dome than he is letting on when he asks him to help get a message to Julia. Back in Chester's Mill, Big Jim appoints himself sheriff of the town.
- The residents of Chester's Mill continue to argue over what to do with the egg to ensure everyone's safety, while Junior is haunted by a familiar face.
- As temperatures continue to plunge, Sam and Rebecca spring into action to try and save the residents of Chester's Mill. Meanwhile, Joe remains suspicious of Hunter, and Barbie and Julia face danger after black ice causes their vehicle to crash.
- When a new threat from the Dome intensifies, the residents of Chester's Mill find themselves at risk of being crushed to death. Meanwhile, Melanie's health continues to deteriorate as the fate of the egg remains unknown.
- A potential exit from the Dome is revealed just as the walls begin closing in on those trapped in Chester's Mill.
- The residents of Chester's Mill appear both inside and outside the dome following their mysterious encounter in the tunnels beneath the town.
- The townspeople are forced to question who they can trust when new residents emerge.
- As the dome comes down, a final attempt is made to keep infected townspeople from impacting the outside world.
- The residents of Chester's Mill try to move on after emerging from the cocoons in the tunnels. Big Jim becomes suspicious of Christine and Eva.
- Under the guise of helping Chester's Mill rebuild, Christine urges the townspeople towards specific individuals and projects that remind them of their experience in the tunnels.